The paper explores working class couples' experiences of female-breadwinning during the Great Recession in Spain. It examines the extent to which couples' adaptations to these gender-atypical work-family arrangements have led to pro- cesses of (un)doing gender. The study is based on the analysis of 24 semi-structured biographical interviews and life history calendars with men and women in 12 heterosexual couples who have gone through different breadwinning statuses during their trajectory. Findings show that men whose partners were primary breadwinners for a period make the greatest effort to preserve the male-breadwinner illusion. In contrast, female breadwinners identify with a co-bread- winner model and do not understate their own economic contribution. Men's insufficient participation in housework and child care is experienced by women with disapproval, which turns into open conflict when the women perform the bulk of such work. The paper concludes that adaptation to unconventional arrangements can constitute a catalyst for processes that undo gender, but more qualitative longitudinal research is needed to determine how economistic and normative factors interact dynamically across different countries and social groups in shaping these processes.